Why Do Refrigerated Tomatoes Turn Mealy After They’re Brought Back to Room Temperature?

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I put ripe tomatoes in the refrigerator to make them last a little longer. After taking them out, though, they sometimes feel soft and grainy rather than juicy, even once they’ve warmed up.

Is that texture change caused by the cold itself, or does it depend on how ripe the tomatoes were when they went into the fridge? I’d also be curious whether different tomato varieties respond differently.

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